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Abstract This study focuses on understanding what drives the previously observed deep nighttime ionospheric hole in the American sector during the January 2013 sudden stratospheric warming (SSW). Performing a set of numerical experiments with the thermosphere‐ionosphere‐mesosphere‐electrodynamics general circulation model (TIME‐GCM) constrained by a high‐altitude version of the Navy Global Environmental Model, we demonstrate that this nighttime ionospheric hole was the result of increased poleward and down magnetic field line plasma motion at low and midlatitudes in response to alteredF‐region neutral meridional winds. Thermospheric meridional wind modifications that produced this nighttime depletion resulted from the well‐known enhancements in semidiurnal tidal amplitudes associated with stratospheric warming (SSWs) in the upper mesosphere and thermosphere. Investigations into other deep nighttime ionospheric depletions and their cause were also considered. Measurements of total electron content from Global Navigation Satellite System receivers and additional constrained TIME‐GCM simulations showed that nighttime ionospheric depletions were also observed on several nights during the January‐February 2010 SSW, which resulted from the same forcing mechanisms as those observed in January 2013. Lastly, the recent January 2021 SSW was examined using Modern‐Era Retrospective Analysis for Research and Applications, Version 2, COSMIC‐2 Global Ionospheric Specification electron density, and ICON Michelson Interferometer for Global High‐Resolution Thermospheric Imaging horizontal wind data and revealed a deep nighttime ionospheric depletion in the American sector was likely driven by modified meridional winds in the thermosphere. The results shown herein highlight the importance of thermospheric winds in driving nighttime ionospheric variability over a wide latitude range. 

Abstract NRLMSIS® 2.0 is an empirical atmospheric model that extends from the ground to the exobase and describes the average observed behavior of temperature, eight species densities, and mass density via a parametric analytic formulation. The model inputs are location, day of year, time of day, solar activity, and geomagnetic activity. NRLMSIS 2.0 is a major, reformulated upgrade of the previous version, NRLMSISE‐00. The model now couples thermospheric species densities to the entire column, via an effective mass profile that transitions each species from the fully mixed region below ~70 km altitude to the diffusively separated region above ~200 km. Other changes include the extension of atomic oxygen down to 50 km and the use of geopotential height as the internal vertical coordinate. We assimilated extensive new lower and middle atmosphere temperature, O, and H data, along with global average thermospheric mass density derived from satellite orbits, and we validated the model against independent samples of these data. In the mesosphere and below, residual biases and standard deviations are considerably lower than NRLMSISE‐00. The new model is warmer in the upper troposphere and cooler in the stratosphere and mesosphere. In the thermosphere, N₂and O densities are lower in NRLMSIS 2.0; otherwise, the NRLMSISE‐00 thermosphere is largely retained. Future advances in thermospheric specification will likely require new in situ mass spectrometer measurements, new techniques for species density measurement between 100 and 200 km, and the reconciliation of systematic biases among thermospheric temperature and composition data sets, including biases attributable to long‐term changes.